Ancient Egyptian Fresco of Scarab Beetle. The adults of most species feed on dung or nectar, or do not feed at all. It might be a Bumble Flower Beetle, Euphoria inda, which is pictured on BugGuide . Connect with your County Extension Office , Find an Extension employee in our staff directory , Get the latest news and updates on Extension's work around the state, Feedback, questions or accessibility issues: info@extension.wisc.edu | 2023 The Board of Regents of the University of Wisconsin System Privacy Policy | Non-Discrimination Policy & How to File a Complaint | Disability Accommodation Requests. They are usually found on flowers, and some species have been observed feeding on aphids and other soft-bodied insects. As with our other list pages, you can click on the small 'X' in each entry to remove unneeded/unwanted entries in the result. Common names tell us about the appearance, toxicity, odor, folklore, uses, habitats and other aspects of organisms as they are observed by us, out here in the field where the rubber hits the road. Several subfamilies have been elevated to family rank (e.g., Bolboceratidae, Geotrupidae, Glaresidae, Glaphyridae, Hybosoridae, Ochodaeidae, and Pleocomidae), and . Ground beetles Family Carabidae, Ground beetles belong to the family Carabidae, which is one of the largest beetle families, with at least 34,000 and maybe more than 40,000 species worldwide. A century ago, in response to this variability, taxonomic splitters ordained 10 species/subspecies, all of which have since been rolled back into todays GB. About the Green June Beetle Chittendon also says that Opinion up to now is that it is an occasional pest, but that opinion is by people who havent studied it. Ninety years later, it seems like some of the answers are still up for grabs. Largest Beetles: Green June Beetle, Japanese Beetle, Black Blister Beetle, Smallest Beetles: Black Carpet Beetle, Khapra Beetle, Varied Carpet Beetle, Most Common Beetles: Japanese Beetle, Varied Carpet Beetle. of Wisconsin-Madison Madison, WI 53706 Office: (608) 262-3229 Fax: (608) 262-3322 groves@entomology.wisc.edu Bio . Really big scarabs in other countries can measure 6 " in length. They also feed on aphids and other insect pests. Encyclopaedia Britannica's editors oversee subject areas in which they have extensive knowledge, whether from years of experience gained by working on that content or via study for an advanced degree. In his monograph Synchronicity (1973) Carl Jung (1875-1961) reported a synchronicity with a patient during psychotherapy. The smaller species, on the other hand, like the carpet and khapra beetles, thrive indoors, destroying carpets, clothes, and food products. X. Scarab Beetles. Pick your answer, all from the literature: Do they mellow out as adults? Some are red or orange with black spots, some are black with red spots, while others have no spots at all. Most adult beetles are nocturnal, although the flower chafers (Cetoniinae) and many leaf chafers (Rutelinae) are active during the day. Updates? Jung commented that the beetle "contrary to its usual habits had evidently felt an urge to get into a dark room at this particular moment." It is also known as spotted pelidnota or spotted June beetle. These products often provide limited and highly variable control compared to chemical insecticides. Some have been shown to be important natural enemies of the eggs and larvae of flies that breed in manure or similar habitats. The grubs mostly live underground or under debris, so are not exposed to sunlight. Unlike most synchronicities, this one had an intermediary (Jung) between the event in the surroundings (the beetle knocking on the window) and the mental event (the patient's dream). Understanding where these diverse beetles fit into the animal kingdom is important because they make up such a big part of the insect world. There wasnt much in between, other than a whole bunch of picture sites. It is pleasant to see how from such a seeming paucity of color, Nature has begotten in this insect such a richness of results., The genus name, Cotalpa, is Latin for with a mole, probably a reference to the fact that this species spends much of its life underground, even as an adult. In the home garden, wireworms cause problems primarily in new vegetable gardens that were formerly lawns. Jung could see that her quest for psychological change was doomed unless he was able to succeed in softening her rationalist shell with a somewhat more human understanding. He needed something to help transform her. https://doi.org/10.1037/h0022912. Though most are not garden ruffians, those that are can cause significant injury. 24. They often pull leaves off of trees as they feed, which annoys homeowners, but they usually live in woodlands where there are plenty of trees. The BugLady found this beetle recently (whats left of it anyway) on what remains of her dune (not much, after the winter storms) (sadly, her Doodlebug Refuge was mostly washed away, and this year houses just a single resident). Several subfamilies have been elevated to family rank (e.g., Bolboceratidae, Geotrupidae, Glaresidae, Glaphyridae, Hybosoridae, Ochodaeidae, and Pleocomidae), and some reduced to lower ranks. Not all beetles have such fanciful names as fireflies and ladybugs. Adult scarabs are generally vegetarians or scavengers. Many species are nocturnal, and many are found associated with the ground, such as under leaf litter or stones, or in loose soil. To rake or not to rake? The six legs, each with five segments (total 30), represent the Larval food sources include fungi, dung, compost and decaying plant material. Most have a bullet-shaped, very hard body. Many scarabs are scavengers that recycle dung, carrion, or decaying plant material. Her amazement triggered positive psychological change. Some scarabs are pretty spectacular https://bugguide.net/node/view/21082, and https://bugguide.net/node/view/73330, and https://bugguide.net/node/view/11561/bgpage. His own actions produced the result he sought. The adults are oval to oval-elongate in shape, but vary a lot in color and shape so they can be difficult to recognize consistently. Scarab Beetles (Scarabaeidae) Scarab beetles are a family of over 30,000 beetles found throughout the world. Dung beetles evolved sixty-five million years ago, as dinosaurs died out and mammals grew larger. As a group, scarabs are nocturnal, oval, chunky, possessed of sturdy front legs that are widened and toothed for digging, and plates at the ends of their antennae (which GBs may hide when theyre sitting still and deploy when theyre moving around). Please refer to the appropriate style manual or other sources if you have any questions. Mollusk feeders live in moist habitats. These beetles are also commonly known as lightning bugs. Habitat? Yet he reported being surprised that the beetle wanted to enter a dark room "contrary to its usual habits.". Ground beetles are swift runners; many species have large mandibles used to hold and dispatch victims. Adult soldier beetles are elongate, soft-bodied beetles similar to fireflies. Check out 8 bugs you should never kill in your garden. Some of the well-known beetles from the Scarabaeidae are Japanese beetles, dung beetles, June beetles, rose chafers (Australian, European, and North American), rhinoceros beetles, Hercules beetles and Goliath beetles. Why Are So Many Young Men Single And Sexless? Scarab-beetle, God with Infant Sun and Sun-Disc below, c11th century BC. Luminescent larvae are often called glow-worms, a name also used for larvae of another family, the Phengodidae. Preventative insecticides must be applied prior to any egg-laying to achieve maximum effectiveness. To rake or not to rake? Some dung beetles form balls of dung that they roll about with their hind legs, sometimes for long distances and sometimes working in pairs. The elongate larvae are covered with tubercles or spines, and usually spotted or banded with bright colors. 2023 The Board of Regents of the University of Wisconsin System, Non-Discrimination Policy & How to File a Complaint. Their larvae, too, are predatory. Beetles undergo complete metamorphosis, meaning that the life stages consist of egg, larva (several instars), pupa, and winged adult. This beetle was associated with the divine manifestation of the early morning sun, Khepri, whose name was written with the scarab hieroglyph and who was believed to roll the disk of the morning sun over the . They probably court via pheromones, and Goldsmith beetles also communicate with sound, rubbing a ridged portion of their body with their legs. Reviewed by Lybi Ma. On this Wikipedia the language links are at the top of the page across from the article title. Fireflies Family Lampyridae, Adult fireflies are medium-sized, dark-colored beetles with soft and flexible elytra. For example, when we here in Gods Country hear the words June Bug, we picture the rusty, shiny, chunky beetle that emerges from the soil in late May and spends the next month buzzing against and clinging on the screens at night, only to disappear underground by sunrise (thats why those hard wing coverselytrawere invented, to protect the soft, flying wings from wear and tear). This fairy tale like coincidence has had major impact not only on the patient and Jung but also on the study of coincidences. If youre unable to find the information you need, please submit your gardening question here: Connecting people with the University of Wisconsin. [5], In Ancient Egypt, the dung beetle now known as Scarabaeus sacer (formerly Ateuchus sacer) was revered as sacred. The strawberry clipper (Anthonomus signatus) damages strawberries by puncturing unopened buds to deposit an egg inside. Chemical Control:Consider using insecticides in turfgrass settings where there are existing or anticipated high grub populations, and in settings (e.g., golf courses, sports fields, etc.) 2023 (Beetle Identifications). Some of the 16,500 species worldwide feed on decaying organic matter while others prefer living plants. This Scarab Beetle is larger than others and can grow to almost 5cm, or 2 inches in length. Some species have similar feeding habits as both larvae and adults, and may even live in the same environment; the aphidophagous (aphid-feeding) lady beetles are one example. A common species in the Midwest is the goldenrod soldier beetle. When numerous, they remove a lot of leaf tissue and can cause significant damage to the plants. Theyre almost twice the size of the common ladybug. In warm tropical climates, lady beetles are active year-round. They are harmless and remind many of the large beetles found in the tropics. They can vary in size from less than inch long to over one inch long depending on the species and their stage of development. This Asian introduction has become naturalized from Maine to South Carolina and west to Wisconsin. Beetle Grub. The scarab story also illustrates a unique variation on a new coincidence form, the meta-coincidence, which will be defined and explored later in this post. Get the help you need from a therapist near youa FREE service from Psychology Today. Feedback, questions or accessibility issues: Click to share on Twitter (Opens in new window), Click to share on Pinterest (Opens in new window), Click to share on Facebook (Opens in new window), Click to share on Tumblr (Opens in new window), Click to share on Reddit (Opens in new window), Click to share on Pocket (Opens in new window), 2023 Board of Regents of the University of Wisconsin System. Adults typically feed on flowers and foliage. At night they come out to feed. Except for the Northwest, the six members of the genus are spread over the whole continent. Note that neonicotinoid insecticides used for grub control can pose risks to bees and other pollinators. Its also called the Fig-eater, a name it shares with a nearly identical scarab (C. mutabilis) that occupies the southwestern corner of the U.S. One source told of children in the South tying strings to GJBs and flying them on their leashes.. The legs are brownish yellow, or brassy, shaded with green. The under part of the insect is a sheet of highly polished copper, from which stands forth a thick coat of whitish wool, justifying its specific name lanigera, wool-bearer. Reinforcement and nonreinforcement in Rogerian psychotherapy. David Mizejewski is a naturalist with the National Wildlife Federation, as well as a nationally recognized media personality and speaker. Current and Future Use of Nematodes in Biocontrol: Practice and Commercial Aspects with Regard to Regulatory Policy Issues. Chronological Index to the Field Station Bulletin, fan out to pick up chemical signals or tuck out of sight, https://bugguide.net/node/view/11561/bgpage, shades of yellow gilded with a metallic sheen, live in woodlands where there are plenty of trees. Famous for their glowing abdomens, many firefly species dont live long as adults because they are too focused on mating and reproducing to eat, but their larvae are predatory, feasting on other insects, slugs and snails. Preventative insecticides provide long-lasting protection. Note: Thank-you to Tara dos Santos for her helpful suggestions. Many species of these stout, oval-shaped beetles feed on animal dung, making them very good bugs for your garden! Both participants were changed by the event. The beetles, order Coleoptera, constitute the single largest group of animals on earth. When disturbed, the adults tuck their legs in and drop to the ground. Following a pheromone-induced romance, Ms. GJB tunnels as deep as 8 underground. Here is the list of beetles found in Wisconsin arranged in order of the family they belong to. They are easily recognized by their elongated snout and elbowed antennae on the snout. For all products, youmustapply approximately inch of water at the time of application to wash the insecticide into the turf root zone where grubs are actively feeding. Larvae feed for a year or two and then pupate in cells they make deep in the earth. The bigger beetles like the Japanese beetle and green june beetle mainly feed on plants like rose, grapes, and sweet potatoes. There are 1,700 scarab species in North America and 28,000 worldwide. Both the adults and larvae are plant feeders. Leaf beetles Family Chrysomelidae. Consult product labels for application instructions and timing. X-number:XHT1018. Adult lady beetles may require other food in addition to prey, and many feed on flower pollen, nectar, or honeydew. There are many destructive species that feed on seeds and roots, and many others are scavengers. Free shipping for many products! In 1868, the Reverend Samuel Lockwood described them more generously: Among the Beetles of North America, very few can bear away the palm for beauty from the Cotalpa lanigera, or, as popularly known, the Goldsmith Beetle. Psychotherapy research suggests that the therapist shapes the content of what patients say. Additional damage can occur when animals (e.g., skunks, raccoons, crows, etc.) Learn how to keep squash bugs out of your garden. The number of aphids that can be consumed by a lady beetle varies somewhat with species of both predator and prey and also with environmental conditions such as temperature. 21. When on their back, these beetles can snap two parts of the body together, spring up into the air, and land on their feet. Although many are black, brownish, or drab, many scarabs are colorful, some with iridescent greens and other colors, others looking as if they were covered with shiny enamel paint. They range in size from 12 to 35 mm (0.47 to 1.38 in) and are blackish or reddish-brown in colour, without prominent markings, and often rather . I always say that instead of using chemicals, its better to have a diversely planted garden that provides lots of habitat for hardworking garden beetles and other pollinators and predators. Applying water maximizes the effectiveness of the insecticide. Treatment timing can vary by white grub species. Are Zoomies a Sign of a Happy Dog or a Crazy Dog? White grubs have pale, C-shaped bodies, orange heads and three pairs of legs. X. The black vine weevil (Otiorhynchus sulcatus) and strawberry root weevil (Otiorhynchus ovatus) are common pests in gardens, with the larvae feeding on the roots of many woody shrubs and strawberry. Of the remaining families which are not mentioned in this article some are primarily aquatic, a few offer natural control of pests of agriculture or forestry, and others are not common. The rose-chafer beetle is common in Central and Southern Europe where Jungs home town of Zurich, Switzerland, is located. In other groups, the larvae and adults do completely different things. Rose chafers are scarab beetles approximately 3/8 inch long, slender, and light tan in color. An egg is laid at the end of each long tunnel and food is left there for the growing grub. Jill Staake Sometimes called dung beetles, scarab beetles are sometimes surprisingly beautiful up close.. Despite their intimidating appearance, these beetles are harmless to people. Im toxic.. These beetles get their common name from their distinctive antennae which are at least the length of the body and sometimes even longer than the body. The adults emerge to feed on the foliage of a variety of ornamental plants. Knowing what you value will help you build the most meaningful life possible. Probably the best known group of beetles, they vary in color and pattern. Beetles occur in virtually all habitable terrestrial and freshwater environments. Jung got up, opened the window, caught the beetle and presented it to her. Insecticide Application Timing by White Grub Species, *Curative treatments are most effective against smaller grubs towards the beginning of the treatment window. The most significant scarab pests in gardens are the chafers or white grubs. It was suggested that as Japanese Beetles spread across the land, GJBs will become more pesty in their wake; Adults gore fruits with a horn on the front of their head to get inside. But many of them have something more important in common: Theyre good bugs for your garden. Although ground beetles are generalist predators, and are frequently abundant, they may not always provide good pest control. They are active July through August. So theres a pretty good chance that youre going to encounter garden beetles. Large C-shaped grubs can be found in rotting stumps and the adults appear in July and August. Clarence Holmes Wildlife/Alamy Stock Photo, pictures that will change how you see bugs, 14 Questions About Hummingbird Feeders Answered by Experts, What to Do if You See a Bald-Faced Hornet Nest, Do not Sell or Share My Personal Information. Connect with your County Extension Office , Find an Extension employee in our staff directory , Get the latest news and updates on Extension's work around the state, Feedback, questions or accessibility issues: info@extension.wisc.edu | 2023 The Board of Regents of the University of Wisconsin System Privacy Policy | Non-Discrimination Policy & How to File a Complaint | Disability Accommodation Requests. Indeed, Madam Cotalpa has long been an acknowledged belle among the Rutilians, themselves a distinguished family in Beetledom. In addition to the aphid predators, there are other species that specialize on other types of prey. The adults are attracted to light and feed on foliage.They can make a hissing sound when touched or otherwise disturbed, which can resemble the hissing of a bat.This sound is made by their wings pushing down, forcing the air out between . In keeping with the food habits of many scarabs, they eat manure and decaying organic matter in the soil and are only minor pests; They eat the roots of grasses especially lawns and golf courses, plus oats, alfalfa, sorghum, tobacco, and some ornamental crops; The larvae cause mechanical damage to plants by uprooting them while feeding, or by creating air spaces that separate roots from the soil moisture, or by loosening them so that livestock pull them up while grazing, leaving unsightly bald spots; Adults are abstemious, mostly foregoing food but sometimes indulging in tree sap or a little moisture from rotted fruits; They drink nectar (especially from open flowers like hollyhocks) and eat ripening fruits (peaches and grapes were most frequently mentioned), as well as corn and the foliage of a great many plants, both wild and tame, on which they can be pests; Adults eat ripe or decaying fruit, in which several may bury themselves at once, fouling it with odor and frass; Because their mandibles are blunt, GJBs cant break into fruits like grapes on their own, but they take advantage of damage initiated by another scarab, the infamous, imported Japanese beetle, that allows them entry (a partnership called facilitationin this case, intra-guild (beetle-to-beetle) facilitation). The BugLady highly recommends the 1868 article by Lockwood for its style and for its insights into the gloriously curious mind that was determined to discover the secrets of the Goldsmith beetle. The adults use their flashing lights in the mating process. Eggs are laid under hay bales in the fields, under moist heaps of manure and compost and mulch, and beneath grass (including pastureland) that has been visited by a manure spreader (either 4-wheeled or 4-footed). Long-horned beetles Family Cerambycidae. Incident #1: The dream image. Omissions? In his monograph Synchronicity (1973) Carl Jung (1875-1961) reported a synchronicity with a patient during psychotherapy.The image of a scarab beetle had appeared in patient's dream and as she . dung beetle, nxai pan national park, botswana - scarab beetle stock pictures, royalty-free photos & images. It consists of both small and large sizes of beetles . An EEO/AA employer, University of Wisconsin-Madison Division of Extension provides equal opportunities in employment and programming, including Title VI, Title IX, the Americans with Disabilities Act (ADA) and Section 504 of the Rehabilitation Act requirements. Many species of these stout, oval-shaped beetles feed on animal dung, making them very good bugs for your garden! If you have goldenrod or Joe Pye weed in your garden, youll very likely see them on those plants. Scarab beetles are one of the most popular families with insect collectors because of the large size and beautifully coloured, hard, highly polished forewings of many species. For more information on white grub control:See University of Wisconsin Garden Facts XHT1062 and XHT1240 (available athttps://pddc.wisc.edu/), University of Wisconsin-Madison Extension bulletins A3275, A3714, A4128, A4130, A4141, A3950, A2303, A3435 and A3434 (available athttps://learningstore.extension.wisc.edu/), the Turf Management Mobile website (https://turfpests.wisc.edu/), or contact your county Extension agent. There is a smaller species that attack raspberries. [1], Scarabs are stout-bodied beetles, many with bright metallic colours, measuring between 1.5 and 160 millimetres (0.059 and 6.3in). The light-colored, grub-like larvae live in the soil, feeding on roots of grasses and other plants for one or more years. Both the adults and larvae feed on plant material. She was describing a dream from the previous night about a costly piece of jewelry in the shape of a golden scarab. Nodjmet was an ancient. X. We teach, learn, lead and serve, connecting people with the University of Wisconsin, and engaging with them in transforming lives and communities. This distinctly colored beetle is common in gardens where milkweed has been allowed to grow. Most lady beetles overwinter in the adult stage and many form overwintering aggregations. The long, white larvae bore into the stems of herbaceous plants or trees, and can be very destructive to trees the introduced Asian, longhorned beetle (Anoplophora glabripennis) is a good example. scarab beetle, (family Scarabaeidae), any of approximately 30,000 species of beetles (insect order Coleoptera) that are compact and heavy-bodied insects with robustly oval outlines. Scarab beetles vary considerably in habits, with many species feeding on manure or on decomposing plant materials, others on growing roots or leaves, and a few on fungi. Legs are thick and have some serration and hairs on them. To have a beetle knocking on Jungs window during spring and early summer would be likely. Lady beetles (more frequently, but less accurately, referred to as ladybugs; also called ladybird beetles) typically have round or oval, strongly convex bodies with bright contrasting colors. However, black and brown are the most common. The wing-cases are a gleaming lemon, thus making the whole back appear as if encased by two large plates of paly gold, while the thorax and head are each covered with brilliant red gold, which in the light gives off an almost flaming hue. Scarabs are part of the Insecta class, Arthropoda phylum, and Animalia kingdom. Their undersides may be shiny black or metallic green, and while the legs of southern GBs are light-colored, Yankee GBs have dark legs. After five years and 244 BOTWs, BugFan Patrick is leaving our midst. The scarab beetle was also associated with the gods Atum and Re, who represented primordial creation and the sun, respectively. The pale, C-shaped larvae hatch two weeks later, dig/bore in, and feedand feed, and feedfor the rest of their first year and through their second summer. For example, almost all ground beetles are predaceous and the leaf beetles are mostly all leaf feeders. Euonymous is one of the plants frequently damaged by adult root weevils. Finally, be sure to properly reseed damaged areas as needed. The species of lady beetles that are aphid predators will occasionally take other types of insects and mites as food. Although some larvae also feed on leaves, many larvae attack plant roots. Many are nocturnal or crepuscular. The common species of lady beetles those that are larger, orange or red, often with black spots are most frequently specialized predators of aphids. Both the larvae and adults of some species are considered pests (the infamous Japanese beetle is a scarab), but other species are decomposers that are important members of their local food webs. When grubs are abundant however, they can cause significant damage leading to irregular brownish patches of turf. Grape pelidnotas can be found throughout the late spring and summer. Learn how to get rid of stink bugs in your garden. When the last larval stage is fully grown, pupation occurs, usually on a plant part, such as foliage or stem, where the larva was when it finished feeding. where tolerance for grub damage is low. In this famous synchronicity, the scarab served as the external event that coincided with the mental events of two people, Jung and his patient. These are not giant ladybugs, nor are they related to ladybugs outside of their mutual beetle-hood. As the name suggests, they are usually found associated with the soil, particularly in damp habitats. Wisconsin has various beetles inhabiting the state, with most of them ill-reputed for infestation and damage to plants, food grains, or household objects. Vanishing Muse: Five years ago, the not-yet-BugLady encountered not-quite-BugFan Patrick and promised to undertake his Insect Education, one bug at a time, and BOTW was born. Large C-shaped grubs can be found in rotting stumps and the adults appear in July and August. The ten-lined June beetle (Polyphylla decemlineata), also known as the watermelon beetle, is a scarab beetle found in the western United States and Canada. Find many great new & used options and get the best deals for EGYPTIAN Pharaonic Stone Faience Beetle Scarab Hiroglyphic at the best online prices at eBay! Proper timing is important to maximize the effectiveness of grub control products. There she deposits up to 20 eggs at a time in a ball of soil she compacts and glues together. These grubs are the larval stages of scarab beetles such as Japanese beetles (see University of Wisconsin Garden Facts XHT1062,Japanese Beetle), May/June beetles (see University of Wisconsin Garden Facts XHT1240,May/June Beetles), European chafers (see University of Wisconsin-Madison Extension bulletin A4141,European Chafer), and northern masked chafers (see University of Wisconsin-Madison Extension bulletin A4130,Northern Masked Chafer). Almost a monochrome, yet the garb of the Goldsmith beetle presents a rich diversity of tints, chiefly of the yellow sorts. If you do not water, the insecticide may provide little or no control. Here are some of the most common beetles in the garden. In the scarab story, one coincidence is that the woman was talking about a scarab and a scarab appeared. He heard a tapping on the window. If you see one, let it be! Lady beetles Family Coccinellidae. This species may or may not have been the inspiration for Poes story The Gold-bug.. Adult masked chafers are golden brown with darker brown . Most of the approximately 5,000 species worldwide (475 in North American) are beneficial predators, but a limited number are phytophagous (plant feeders) the Mexican bean beetle (Epilachna varivestis) is the most notable. The four most important families of beetles with common garden pests and two additional families that have some significant pests of ornamental plants are reviewed here. As a group, scarabs are nocturnal, oval, chunky, possessed of sturdy front legs that are widened and toothed for digging, and "plates" at the ends of their antennae (which GBs may hide when they're sitting still and deploy when they're moving around). In the upper Midwest, there are generally 2-3 generations per year. The adults also feed on foliage, producing characteristic notches along leaf margins, that is often mistaken for grasshopper feeding. Most turfgrass areas in Wisconsin have white grubs but typically in low numbers that cause minimal damage. Like any other insect out there, the scarab beetles come in a lot of different colors, shades, and hues that may include dark green, vibrant orange-ish, purple, or maybe even rainbow!. Sunflower Tortoise Beetle Larva. The largest of beetle families, with about 48,000 species worldwide, is comprised almost entirely of plant feeders, either on plants or stored grains.

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